1
THE TROPICAL RAINFOREST: HISTORY AND ENVIRONMENT
This is the forest primeval
Henry Longfellow, Evangeline (1847)
1.1 INTRODUCTION
Tropical rainforests are the most complex ecosystems on earth. Rainforests (better known to many people as jungles) have been the dominant form of vegetation in the tropics for literally millions of years, and beneath their high canopy lives a diversity of species which is unrivalled anywhere else on earth.
1.1a Images and impressions
For more than a century travellers have recorded vivid descriptions of the rainforests, which emphasise abundance and grandeur. Charles Darwin kept a detailed log of his impressions of the forests around Rio de Janeiro, which he visited in April 1832 during the voyage of the Beagle. He wrote:
After passing through some cultivated country, we entered a forest, which in the grandeur of all its parts could not be exceeded. âŚThe trees were very lofty, and remarkable, compared with those of Europe, from the whiteness of their trunksâŚ. The forest abounded with beautiful objectsâŚ. The greater number of trees, although so lofty, are not more than three or four feet in circumferenceâŚ. It is easy to specify the individual objects of admiration in these grand scenes; but it is not possible to give an adequate idea of the higher feelings of wonder, astonishment, and devotion, which fill and elevate the mind.1
Since about 1980 interest in the rainforest has shifted full circle. From being seen as a threat or nuisance, it is now widely seen as under threat, with mounting concern for its future survival. The place to be tamed and conquered is how viewed as a place to preserve and protect.
This about-turn in attitude has been triggered by the real threat of destruction and clearance of the rainforest habitat world-wide. Although clearance per se is not new, the pace of deforestation is faster than ever before, and there are real fears that within one generation there will be hardly any natural rainforest left anywhere in the world.
There is already a fairly sizeable literature on the character and dynamics of tropical rainforests.2 In this book we will examine why the rainforest is so important, why it is being cleared, what the consequences of this clearance are and what solutions are available. But first we need to define exactly what we mean by the term ârainforestâ, examine where it is found and reflect on why it is regarded as the most important ecosystem on earth.
1.1b Classification
The rainforest is one of several types of forest found throughout the tropics, and each type has different characteristics.3
The closed forests account for about half of the total area of tropical forest (around 62 per cent of the natural tropical forest) and comprise two types of continuous tree cover (Table 1.1). Eleven-twelfths of the closed forests, by area, are tropical moist forests and the rest are deciduous and semi-deciduous forests of various types. About two-thirds of the moist forests are tropical rainforests, composed of evergreen broadleaved trees which flourish in the high temperature and humidity of the low latitudes. The tropical moist deciduous forests (or monsoon forests) grow on the fringes of the tropical rainforests, and lose their leaves in the dry season.
Table 1.1 Distribution of tropical forest types
Most of the remaining tropical forests are open woodland, including shrublands and types of savanna, pasture and grassland which are partly wooded.
Almost all (97 per cent) of the tropical forests which have been modified by human activity are fallow forests, areas which have recently been farmed and then abandoned or left to regenerate naturally. Only a very small area is covered by tropical forest plantations. The industrial plantations produce commercial timber, pulpwood or charcoal; the non-industrial plantations are mainly for fuelwood production or environmental protection.
1.1c Distribution
The tropical rainforests provide a discontinuous belt of green around the globe, between the tropic of Cancer (23.5° north) and the tropic of Capricorn (23.5° south). Dense rainforest is the natural climax vegetation of the hot, humid tropical zone and it flourishes particularly in the lower latitudes (between 10° north and south of the equator). Just under half of the tropical zone (49 per cent according to the World Resources Institute)4 is covered by forests (Figure 1.1).
Most of the tropical countries with surviving rainforests are developing countries, for whom the forests provide a valuable capital asset.
The total area presently covered by tropical rainforests is estimated at 12 million km2, which accounts for nearly a third of the worldâs forests (covering roughly 30 million km2).5 The distribution of forests within the tropics is uneven, reflecting the distribution of land and sea and the impacts of this on climatic boundaries. The latitudinal boundaries of the rainforest are determined mainly by precipitation, while altitudinal limits are determined more by temperature. Some rainforests thrive beyond the 10° north and south latitudes, where high rainfall encourages forest growth. Such patches occur in Central America, the north-east coast of Australia and the great valleys of southern China.
The main rainforests today are found in three areas (Figure 1.1 and Table 1.2)âLatin America, Western Equatorial Africa and South-East Asia. Latin America houses the American Formation which is dominated by the Amazon and Orinoco Basins. It has over half (56 per cent) of the world total, much of it (3.31 million km2, 48 per cent of the areaâs total) in Brazil and the rest in Peru, Ecuador, Colombia, Venezuela and French Guiana. Amazonia is the worldâs largest and most important surviving rainforest.6
The remaining rainforests are scattered in sixteen countries in West and Central Africa (18 per cent of the world total) and South-East Asia (25 per cent of the world total). The African Formation includes the Cameroons and the Congo Basin in countries such as Gabon, ZaĂŻre and Madagascar. The Indo-Malaysian Formation in South-East Asia includes parts of western and southern India, the Far East (especially in Indonesiaâparticularly Borneo and Papua New Guineaâwhich now has about 10 per cent of the worldâs remaining tropical rainforest) and north Australia.
Figure 1.1Global distribution of tropical forests.
Source: after Gross (1990)
Table 1.2 Distribution of tropical forests
1.1d Lack of reliable information
It is surprisingly difficult to define exactly what the total area of rainforest is today, for various reasons.7 Some countries have better (and more complete) survey coverage than others, and not all of the figures available refer to the same year. Recent developments in remote sensing technologies promise much better survey coverage in the future,8 although between 1970 and 1988 less than 60 per cent of tropical forests had been surveyed using any form of remote sensing, particularly available satellite technologies which are costly.9
Figures reported by the United Nations Food and Agriculture Organisation and by individual countries are often underestimates. Figures quoted by campaigning groups must also be treated with some caution, because such groups may well overestimate rates of forest clearance and underestimate the size of remaining areas. Estimates also vary because they include different types of forest cover. Some, for example, include coniferous and bamboo forests as well as broadleaved forests, whereas others do not. It is also difficult to distinguish between primary (original) and secondary (regrowth) forest cover, even in satellite images, so that quoted figures might not always be comparing like with like.
The most recent reliable information on areas of tropical forests comes from the World Resources Institute.10 These figures (Table 1.2) provide a valuable framework for examining the distribution of tropical forests today and a useful baseline for assessing rates and patterns of deforestation (in Chapter 2).
The proportion of the earthâs land surface covered by rainforest is small (around 8 per cent),11 but its significance is considerable. Possibly half (up to 90 per cent according to some experts) of the total number of species of plants and animals found anywhere on earth are found either exclusively or mainly within tropical forests.12 We will evaluate the wider significance of the rainforests in Chapter 4, particularly in terms of what would be lost if all remaining forests were to be cleared.
1.2 CLIMATE AND RAINFORESTS
Climate exerts a strong influence over the broad distribution of rainforests within the tropics. It also has a marked effect on regional patterns and structures of rainforest vegetation and habitat.
Rainforests occur in hot moist climates. They receive more solar radiation throughout the year than any other vegetation zone on earth, which promotes the rich variety and luxurious character of vegetation.
1.2a Characteristic climate
The typical rainforest climate has two main distinguishing featuresârelatively constant temperatures, and heavy rainfall. Tropical forests grow under a narrow range of temperatures but a fairly wide range of precipitation (Figure 1.2). The combination of these two climatic controls creates the very special environment for rainforest growth.
Temperatures in rainforest areas are high, and they vary relatively little throughout the year. Temperatures remain fairly constant at between 20°C and 28°C through the year,13 the warmest months perhaps a degree or so higher than the coldest months in a given place. This uniformity occurs because the sun is mostly overhead, so variations in the length of daylight throughout the year are limited.
Temperature variations are often greater from day to night than from month to month, so that rainforest dynamics are more variable over short-term diurnal cycles than over longer seasonal or annual timescales. Diurnal temperature variations might be as high as 17°C. Intense heat during the afternoon (when temperatures may reach 35°C) gives way to cold at night (temperatures may fall as low as 18°C), then fresh conditions in the morning, in a never-ending rhythm of daily change. Many tropical areas have a fairly predictable daily cycle of weather dictated by temperature and ...